SAN FRANCISCO: The famed gorilla Koko, known for mastering sign language and building a rapport with humans, has died in California, a research center has announced. She was 46.

Born Hanabi-ko (Japanese for "Fireworks Child") on US independence day in 1971 at San Francisco Zoo, she was widely known by her nickname. Animal psychologist Penny Patterson began teaching Koko sign language in 1972.

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