MANILA: Bertha, believed to be the world's oldest hippopotamus, has died aged 65, the Manila zoo said Monday, having beaten the typical lifespan for the mostly herbivorous mammals by decades.

The 2.5-ton female was found dead Friday in her enclosure, with a post mortem examination concluding that Bertha, the zoo's oldest resident, had died from multiple organ failure, zoo director James Dichaves said.

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